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Sub-site nomenclature

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A nomenclature for enzymatic sub-sites of carbohydrate processing enzymes was proposed in 1997 [1]. Sub-sites are numbered with increasingly negative numbers (-1, -2, -3, etc) away from the cleavage point towards the non-reducing terminus, and with increasingly positive numbers (+1, +2, +3, etc) towards the reducing terminus as shown below.

Subsite nomenclature.png


  1. Davies GJ, Wilson KS, and Henrissat B. (1997) Nomenclature for sugar-binding subsites in glycosyl hydrolases. Biochem J. 321 ( Pt 2), 557-9. DOI:10.1042/bj3210557 | PubMed ID:9020895 | HubMed [Davies1997]